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They are the smallest of birds, most species measuring 7. They are known as hummingbirds because of the humming sound created by their beating wings , which flap at high frequencies audible to humans.
They hover in mid-air at rapid wing-flapping rates, which vary from around 12 beats per second in the largest species, to in excess of 80 in some of the smallest.
Hummingbirds have the highest mass-specific metabolic rate of any homeothermic animal. While all hummingbirds depend on flower nectar to fuel their high metabolisms and hovering flight, coordinated changes in flower and bill shape stimulated the formation of new species of hummingbirds and plants.
Due to this exceptional evolutionary pattern, as many as hummingbird species can coexist in a specific region, such as the Andes range.
The hummingbird evolutionary tree shows ancestral hummingbirds splitting from insectivorous swifts family Apodidae and treeswifts family Hemiprocnidae about 42 million years ago, probably in Eurasia.
The Andes Mountains appear to be a particularly rich environment for hummingbird evolution because diversification occurred simultaneously with mountain uplift over the past 10 million years.
Within the same geographic region, hummingbird clades co-evolved with nectar-bearing plant clades, affecting mechanisms of pollination.
Hummingbirds exhibit sexual size dimorphism according to Rensch's rule ,  in which males are smaller than females in small species, and males are larger than females in large-bodied species.
Sexual size and bill differences likely evolved due to constraints imposed by courtship because mating displays of male hummingbirds require complex aerial maneuvers.
Female hummingbirds tend to be larger, requiring more energy, with longer beaks that allow for more effective reach into crevices of tall flowers for nectar.
Another evolutionary cause of this sexual bill dimorphism is that the selective forces from competition for nectar between the sexes of each species are what drive the sexual dimorphism.
Hummingbirds are specialized nectarivores  and are tied to the ornithophilous flowers upon which they feed. This coevolution implies that morphological traits of hummingbirds, such as bill length, bill curvature, and body mass are correlated with morphological traits of plants, for example corolla length, curvature, and volume.
However, even in the most specialized hummingbird-plant mutualisms the number of food plant lineages of the individual hummingbird species increases with time.
Many plants pollinated by hummingbirds produce flowers in shades of red, orange, and bright pink, though the birds will take nectar from flowers of other colors as well.
Hummingbirds can see wavelengths into the near-ultraviolet, but hummingbird-pollinated flowers do not reflect these wavelengths as many insect-pollinated flowers do.
This narrow color spectrum may render hummingbird-pollinated flowers relatively inconspicuous to most insects , thereby reducing nectar robbing. Upon maturity, males of a particular species, Phaethornis longirostris, the long-billed hermit , appear to be evolving a daggerlike weapon on the beak tip as a secondary sexual trait to defend mating areas.
In traditional taxonomy , hummingbirds are placed in the order Apodiformes , which also contains the swifts. However, some taxonomists have separated them into their own order, the Trochiliformes.
Hummingbirds' wing bones are hollow and fragile, making fossilization difficult and leaving their evolutionary history poorly documented.
Though scientists theorize that hummingbirds originated in South America, where species diversity is greatest, possible ancestors of extant hummingbirds may have lived in parts of Europe to what is southern Russia today.
Between and species of hummingbirds are described, depending on taxonomic viewpoint, divided into two subfamilies , the hermits subfamily Phaethornithinae, 34 species in six genera , and the typical hummingbirds subfamily Trochilinae , all the others.
However, recent phylogenetic analyses suggest that this division is slightly inaccurate, and that there are nine major clades of hummingbirds: the topazes and jacobins , the hermits, the mangoes , the coquettes, the brilliants, the giant hummingbird Patagona gigas , the mountaingems , the bees, and the emeralds.
The hummingbird family has the second-greatest number of species of any bird family after the tyrant flycatchers. Fossil hummingbirds are known from the Pleistocene of Brazil and the Bahamas ; however, neither has yet been scientifically described, and fossils and subfossils of a few extant species are known.
Until recently, older fossils had not been securely identifiable as those of hummingbirds. In , Dr. Gerald Mayr of the Senckenberg Museum in Frankfurt am Main identified two million-year-old hummingbird fossils.
Fossils of birds not clearly assignable to either hummingbirds or a related, extinct family, the Jungornithidae , have been found at the Messel pit and in the Caucasus , dating from 40—35 mya ; this indicates that the split between these two lineages indeed occurred at that date.
The areas where these early fossils have been found had a climate quite similar to that of the northern Caribbean or southernmost China during that time.
The biggest remaining mystery at the present time is what happened to hummingbirds in the roughly 25 million years between the primitive Eurotrochilus and the modern fossils.
The astounding morphological adaptations , the decrease in size, and the dispersal to the Americas and extinction in Eurasia all occurred during this timespan.
DNA-DNA hybridization results  suggest that the main radiation of South American hummingbirds took place at least partly in the Miocene , some 12 to 13 million years ago, during the uplifting of the northern Andes.
In , a million-year-old bird fossil unearthed in Wyoming was found to be a predecessor to both hummingbirds and swifts before the groups diverged.
The highest recorded wingbeats for wild hummingbirds during hovering is 88 per second, as measured for the purple-throated woodstar Calliphlox mitchellii weighing 3.
During turbulent airflow conditions created experimentally in a wind tunnel , hummingbirds exhibit stable head positions and orientation when they hover at a feeder.
During evolution, hummingbirds have adapted to the navigational needs of visual processing while in rapid flight or hovering by development of an exceptionally dense array of retinal neurons allowing for increased spatial resolution in the lateral and frontal visual fields.
Hummingbirds are highly sensitive to stimuli in their visual fields, responding to even minimal motion in any direction by reorienting themselves in midflight.
With the exception of insects , hummingbirds while in flight have the highest metabolism of all animals — a necessity to support the rapid beating of their wings during hovering and fast forward flight.
Hummingbirds can use newly-ingested sugars to fuel hovering flight within 30—45 minutes of consumption. A review indicated that hummingbirds have in their flight muscles a mechanism for "direct oxidation" of sugars into maximal ATP yield to support their high metabolic rate for hovering, foraging at altitude, and migrating.
By relying on newly-ingested sugars to fuel flight, hummingbirds can reserve their limited fat stores to sustain their overnight fasting or to power migratory flights.
The high metabolic rate of hummingbirds — especially during rapid forward flight and hovering — produces increased body heat that requires specialized mechanisms of thermoregulation for heat dissipation, which becomes an even greater challenge in hot, humid climates.
While hovering, hummingbirds do not benefit from the heat loss by air convection during forward flight, except for air movement generated by their rapid wing-beat, possibly aiding convective heat loss from the extended feet.
The dynamic range of metabolic rates in hummingbirds  requires a parallel dynamic range in kidney function. Hummingbird kidneys also have a unique ability to control the levels of electrolytes after consuming nectars with high amounts of sodium and chloride or none, indicating that kidney and glomerular structures must be highly specialized for variations in nectar mineral quality.
Consisting of chirps, squeaks, whistles and buzzes,  hummingbird songs originate from at least seven specialized nuclei in the forebrain.
The metabolism of hummingbirds can slow at night or at any time when food is not readily available: the birds enter a hibernatory, deep-sleep state known as torpor to prevent energy reserves from falling to a critical level.
During torpor, to prevent dehydration , the GFR Glomerular Filtration Rate ceases, preserving needed compounds such as glucose, water, and nutrients.
Use and duration of torpor vary among hummingbird species and are affected by whether a dominant bird defends territory, with nonterritorial subordinate birds having longer periods of torpor.
Hummingbirds have unusually long lifespans for organisms with such rapid metabolisms. Though many die during their first year of life, especially in the vulnerable period between hatching and fledging , those that survive may occasionally live a decade or more.
As far as is known, male hummingbirds do not take part in nesting. Many hummingbird species use spider silk and lichen to bind the nest material together and secure the structure.
Hummingbird building a nest in San Diego Zoo. Video Clip. Nest with two nestlings in Santa Monica, California. Feeding two nestlings in Grand Teton National Park.
Fallen Anna's hummingbird nest in Ventura County, California , shown next to a toothpick for scale. To serve courtship and territorial competition , many male hummingbirds have plumage with bright, varied coloration  resulting both from pigmentation in the feathers and from prismal cells within the top layers of feathers of the head, gorget , breast, back and wings.
By merely shifting position, feather regions of a muted-looking bird can instantly become fiery red or vivid green.
One study of Anna's hummingbirds found that dietary protein was an influential factor in feather color, as birds receiving more protein grew significantly more colorful crown feathers than those fed a low-protein diet.
Hummingbird flight has been studied intensively from an aerodynamic perspective using wind tunnels and high-speed video cameras.
Two studies of rufous or Anna's hummingbirds in a wind tunnel used particle image velocimetry techniques to investigate the lift generated on the bird's upstroke and downstroke.
Many earlier studies had assumed that lift was generated equally during the two phases of the wingbeat cycle, as is the case of insects of a similar size.
Because of their flying technique, these birds no longer have an alula , while the alula digit has evolved to become absent. The giant hummingbird's wings beat as few as 12 beats per second  and the wings of typical hummingbirds beat up to 80 times per second.
A slow-motion video has shown how the hummingbirds deal with rain when they are flying. To remove the water from their heads, they shake their heads and bodies, similar to a dog shaking, to shed water.
For instance, it is about twice the diving speed of peregrine falcons in pursuit of prey. Note: G-force is generated as the bird pulls out of the dive.
The outer tail feathers of male Anna's Calypte anna and Selasphorus hummingbirds e. Many other species of hummingbirds also produce sounds with their wings or tails while flying, hovering or diving, including the wings of the calliope hummingbird ,  broad-tailed hummingbird , rufous hummingbird , Allen's hummingbird , and streamertail , as well as the tail of the Costa's hummingbird and the black-chinned hummingbird , and a number of related species.
Male rufous and broad-tailed hummingbirds genus Selasphorus have a distinctive wing feature during normal flight that sounds like jingling or a buzzing shrill whistle.
Behaviorally, the trill serves several purposes: . Hummingbirds are restricted to the Americas from south central Alaska to Tierra del Fuego , including the Caribbean.
The greatest species richness is in humid tropical and subtropical forests of the northern Andes and adjacent foothills, but the number of species found in the Atlantic Forest , Central America or southern Mexico also far exceeds the number found in southern South America, the Caribbean islands, the United States, and Canada.
While fewer than 25 different species of hummingbirds have been recorded from the United States and fewer than 10 from Canada and Chile each,  Colombia alone has more than  and the comparably small Ecuador has about species.
The migratory ruby-throated hummingbird breeds in a range from the southeastern United States to Ontario ,  while the black-chinned hummingbird , its close relative and another migrant, is the most widespread and common species in the southwestern United States.
The rufous hummingbird is the most widespread species in western North America,  and the only hummingbird to be recorded outside of the Americas, having occurred in the Chukchi Peninsula of Russia.
A few species are year-round residents of Florida , California, and the far southwestern desert regions of the US. Ruby-throated hummingbirds are common along the Atlantic flyway and migrate in summer from as far north as Atlantic Canada ,  returning to Mexico , South America, southern Texas, and Florida to winter.
The rufous hummingbird breeds farther north than any other species of hummingbird,  often breeding in large numbers in temperate North America and wintering in increasing numbers along the coasts of the subtropical Gulf of Mexico and Florida, rather than in western or central Mexico.
This cold hardiness enables it to survive temperatures below freezing, provided that adequate shelter and food are available.
As calculated by displacement of body size, the rufous hummingbird makes perhaps the longest migratory journey of any bird in the world.
At just over 3 in long, rufous birds travel 3, miles one-way from Alaska to Mexico in late summer, a distance equal to 78,, body lengths.
The northward migration of rufous hummingbirds occurs along the Pacific flyway  and may be time-coordinated with flower and tree leaf emergence in spring in early March, and also with availability of insects as food.
For nutrition , hummingbirds eat a variety of insects, including mosquitoes , fruit flies , gnats in flight or aphids on leaves and spiders in their webs.
To supply energy needs, hummingbirds drink nectar, a sweet liquid inside certain flowers. Nectar is a mixture of glucose, fructose, and sucrose, and is a poor source of nutrients , requiring hummingbirds to meet their nutritional needs by consuming insects.
Hummingbirds do not spend all day flying, as the energy cost would be prohibitive; the majority of their activity consists simply of sitting or perching.
Because their high metabolism makes them vulnerable to starvation , hummingbirds are highly attuned to food sources. Some species, including many found in North America, are territorial and will try to guard food sources such as a feeder against other hummingbirds, attempting to ensure a future food supply for itself.
Additionally, hummingbirds have an enlarged hippocampus , a brain region facilitating spatial memory used to map flowers previously visited during nectar foraging.
Hummingbird beaks are flexible  and their shapes vary dramatically as an adaptation for specialized feeding.
Thornbills have short, sharp bills adapted for feeding from flowers with short corollae and piercing the bases of longer ones.
The sicklebills' extremely decurved bills are adapted to extracting nectar from the curved corollae of flowers in the family Gesneriaceae.
The bill of the fiery-tailed awlbill has an upturned tip, as in the avocets. The male tooth-billed hummingbird has barracudalike spikes at the tip of its long, straight bill.
The two halves of a hummingbird's bill have a pronounced overlap, with the lower half mandible fitting tightly inside the upper half maxilla. When a hummingbird feeds on nectar, the bill is usually opened only slightly, allowing the tongue to dart out and into the interior of flowers.
Perception of sweetness in nectar evolved in hummingbirds during their genetic divergence from insectivorous swifts , their closest bird relatives.
Hummingbirds drink with their long tongues by rapidly lapping nectar. Their tongues have tubes which run down their lengths and help the hummingbirds drink the nectar.
Such feeders allow people to observe and enjoy hummingbirds up close while providing the birds with a reliable source of energy, especially when flower blossoms are less abundant.
A negative aspect of artificial feeders, however, is that the birds may seek less flower nectar for food, so they reduce the amount of pollination their feeding naturally provides.
White granulated sugar is the best sweetener to use in hummingbird feeders. Red food dye was once thought to be a favorable ingredient for homemade solutions, but it is unnecessary, and there is no point in adding it to the nectar.
Some people speculate red dye could be bad for the birds, although this claim has not received scientific attention. Other animals also visit hummingbird feeders.
Bees, wasps , and ants are attracted to the sugar-water and may crawl into the feeder, where they may become trapped and drown. Orioles , woodpeckers , bananaquits , raccoons and other larger animals are known to drink from hummingbird feeders, sometimes tipping them and draining the liquid.
Hummingbirds have exceptional visual acuity providing them with discrimination of food sources while foraging. In at least one hummingbird species — the green-backed firecrown Sephanoides sephaniodes — flower colors preferred are in the red-green wavelength for the bird's visual system , providing a higher contrast than for other flower colors.
Some species of sunbirds of Africa, southern and southeastern Asia, and Australia resemble hummingbirds in appearance and behavior, as do perhaps also the honeyeaters of Australia and Pacific islands.
These two groups, however, are not related to hummingbirds, as their resemblance is due to convergent evolution.
The hummingbird moth is often mistaken for a hummingbird. Hummingbird feeding from a flower in the University of California Botanical Garden.
From Wikipedia, the free encyclopedia. For other uses, see Hummingbird disambiguation. Family of birds. Play media. See also: List of Apodiformes by population.
Hummingbird with yellow pollen on its beak in the University of California Botanical Garden. Talamanca hummingbird. Costa's hummingbird. Birds portal Animals portal Biology portal.
The Birds of Ecuador, Field Guide 1 ed. Cornell University Press. Archived from the original on Retrieved Retrieved 30 September Systematic Biology.
Current Biology. Remsen; Dudley, R. Journal of Ornithology. Bibcode : Sci BMC Evolutionary Biology. Biological Journal of the Linnean Society. The American Naturalist.
De; Rowe, Locke Evolutionary Biology. Annals of the Missouri Botanical Garden. Morphological traits determine specialization and resource use in plant—hummingbird networks in the neotropics.
Ecology, 95 12 , Journal of Biogeography. Retrieved 8 March Bibcode : Oecol. PLOS Biol. Plant Syst. A test for sexually dimorphic weapons in a lekking hummingbird".
Behavioral Ecology. Stave off inanition with the word morsels from this month! Words nearby humming Humism , humiture , hummable , hummel , hummer , humming , hummingbird , hummingbird moth , humming top , hummock , hummus.
Words related to humming busy , brisk , buzzing , bustling , hopping. Example sentences from the Web for humming It starts off like any other Lana tune, replete with minor chords and humming , distorted vocals.
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Example sentences from the Web for humming It starts off like any other Lana tune, replete with minor chords and humming , distorted vocals.
The Silly Syclopedia Noah Lott. The Open Air Richard Jefferies. Methods of Authors Hugo Erichsen. Amaryllis at the Fair Richard Jefferies.
Free Grammar Check! Current Zool. J Exp Biol. Proceedings of the National Academy of Sciences. Bibcode : PNAS.. Brain Behav Evol.
J Comp Neurol. Retrieved 24 April The Journal of Experimental Biology. Physiological Zoology. Nutrition Journal. Journal of Comparative Physiology B.
Functional Ecology. Journal of Experimental Biology. The Life of the Hummingbird. New York: Crown Publishers. Royal Society Open Science. Physiological and Biochemical Zoology.
Mental Floss. Retrieved 14 January Comparative Biochemistry and Physiology. Part A. Journal of Avian Biology. Journal of Morphology.
All About Birds. Retrieved 25 June Bibcode : Natur. Journal of Comparative Physiology A. General and Comparative Endocrinology. The natural history of shrews.
Global Ecology and Biogeography. Bulletin of the Texas Ornithological Society. Public Broadcasting System — Nature; from Learner.
Archived from the original video on 2 February Retrieved 12 May Retrieved 12 May — via YouTube. Retrieved 21 June Sciencing, Leaf Group Media.
Retrieved 8 February Annenberg Learner, The Annenberg Foundation. Section: Plumage and Molt. Houghton Mifflin Harcourt.
The Wilson Bulletin. Avian Flight. Oxford University Press, Ornithology Series. BBC News. Retrieved 1 Sep Retrieved March 22, Fleur N July 20, Huffington Post.
Aviation, Space, and Environmental Medicine. Bibcode : PLoSO An apparent example of sexual sensory bias". Current Zoology.
Cosmos Magazine. Animal Behaviour. Retrieved 13 July — via hummingbirds. Heynen Genus Oreotrochilus. Handbook of the Birds of the World.
Barn-owls to Hummingbirds. Lynx Edicions, Barcelona. Barros Species lists of birds for South American countries and territories: Chile.
Caro Checklist to the Birds of Colombia Archived at the Wayback Machine Conservation Colombiana 8. Species lists of birds for South American countries and territories: Ecuador.
The Ontario Hummingbird Project. Archived from the original on 20 April Retrieved 3 May Houghton Mifflin Co. Retrieved 5 Feb Hummingbird Central.
Retrieved 28 August Archived from the original on April 3, Retrieved March 23, Cornell University Laboratory of Ornithology.
Retrieved 10 April Journey North, Annenberg Learner, learner. Archived from the original on 7 March Retrieved 22 March Retrieved 10 May Cornell University, Laboratory of Ornithology, Allaboutbirds.
Archived from the original on 16 July Retrieved 24 January Princeton University Press. Edward; Hume, Ian D. Comparative Physiology of the Vertebrate Digestive System.
Cambridge University Press. Biology Letters. Dead link 2. Evolution of sweet taste perception in hummingbirds by transformation of the ancestral umami receptor".
Am J Clin Nutr. May 2, The New York Times. Annals of Botany. Ecology and Evolution. Audubon Society. Audubon California Kern River Preserve.
Retrieved on Attracting and Feeding Hummingbirds. Wild Birds Series T. The Firefly Forest. Humming without realizing AdamA.
My brothers got this thing where he'll hum and not realize it. It's pretty loud, and it's not like he's humming a tune or anything. It's just one loud long note.
We'll tell him to stop, he'll say sorry, and then 5 seconds later he could start up again. It doesn't happen all the time.
It seems to happen more when he's by himself, like on the computer or something. It happened once when he was younger, but we got rid of it over a summer, and now its back.
Read 5 Responses. Follow - Does anyone know what the 'technical' term, for this is at least or at least a better description then humming without realizing?
I've tried doing a little research about it on google, etc I don't understand why this is a problem. Isn't this just a habit?
I hum without realizing. My friends and family will tell me so. I don't realize it when I do it. I don't think it is terribly loud and I don't think I am offending anyone.
They tease me about it because they think it is somewhat odd but I really don't think it bothers them. When they tease me, I think about it and try not to do it.
Like your brother, I too suffer with incessant humming.